diff --git a/.github/workflows/test.yml b/.github/workflows/test.yml
index 28c29cc9b..d760c6e74 100644
--- a/.github/workflows/test.yml
+++ b/.github/workflows/test.yml
@@ -13,9 +13,8 @@ jobs:
     strategy:
       matrix:
         node_version:
-          - 14
-          - 16
           - 18
+          - 20
     steps:
       - uses: actions/checkout@v3
       - name: Use Node.js ${{ matrix.node_version }}
diff --git a/README.md b/README.md
index aab44ccb5..899312399 100644
--- a/README.md
+++ b/README.md
@@ -324,6 +324,7 @@ Middleware for Node's built in http server or [`express`](https://expressjs.com/
 
 ```js
 const { App, createNodeMiddleware } = require("@octokit/app");
+
 const app = new App({
   appId: 123,
   privateKey: "-----BEGIN PRIVATE KEY-----\n...",
@@ -337,11 +338,20 @@ const app = new App({
 });
 
 const middleware = createNodeMiddleware(app);
-
-require("http").createServer(middleware).listen(3000);
+require("http")
+  .createServer(async (req, res) => {
+    // `middleware` returns `false` when `req` is unhandled (beyond `/api/github`)
+    if (await middleware(req, res)) return;
+    res.writeHead(404);
+    res.end();
+  })
+  .listen(3000);
 // can now receive user authorization callbacks at /api/github/*
 ```
 
+The middleware returned from `createNodeMiddleware` can also serve as an
+`Express.js` middleware directly.
+
 <table width="100%">
   <thead align=left>
     <tr>
@@ -392,33 +402,9 @@ All exposed paths will be prefixed with the provided prefix. Defaults to `"/api/
 
 Used for internal logging. Defaults to [`console`](https://developer.mozilla.org/en-US/docs/Web/API/console) with `debug` and `info` doing nothing.
 
-</td>
+      </td>
     </tr>
-    <tr>
-      <th>
-        <code>options.onUnhandledRequest</code>
-      </th>
-      <th>
-        <code>function</code>
-      </th>
-      <td>
-
-Defaults to
 
-```js
-function onUnhandledRequest(request, response) {
-  response.writeHead(400, {
-    "content-type": "application/json",
-  });
-  response.end(
-    JSON.stringify({
-      error: error.message,
-    }),
-  );
-}
-```
-
-</td></tr>
   </tbody>
 </table>
 
diff --git a/package-lock.json b/package-lock.json
index f1f1425d6..802ab9600 100644
--- a/package-lock.json
+++ b/package-lock.json
@@ -9,33 +9,31 @@
       "version": "0.0.0-development",
       "license": "MIT",
       "dependencies": {
-        "@octokit/auth-app": "^4.0.13",
-        "@octokit/auth-unauthenticated": "^3.0.0",
-        "@octokit/core": "^4.0.0",
-        "@octokit/oauth-app": "^4.0.7",
-        "@octokit/plugin-paginate-rest": "^6.0.0",
-        "@octokit/types": "^9.0.0",
-        "@octokit/webhooks": "^10.0.0"
+        "@octokit/auth-app": "^6.0.0",
+        "@octokit/auth-unauthenticated": "^5.0.0",
+        "@octokit/core": "^5.0.0",
+        "@octokit/oauth-app": "^6.0.0",
+        "@octokit/plugin-paginate-rest": "^8.0.0",
+        "@octokit/types": "^11.1.0",
+        "@octokit/webhooks": "^12.0.1"
       },
       "devDependencies": {
         "@octokit/tsconfig": "^2.0.0",
         "@types/jest": "^29.0.0",
         "@types/node": "^18.0.0",
-        "@types/node-fetch": "^2.5.8",
         "esbuild": "^0.18.0",
         "express": "^4.17.1",
         "fetch-mock": "^9.10.7",
         "glob": "^10.0.0",
         "jest": "^29.0.0",
         "mockdate": "^3.0.2",
-        "node-fetch": "^2.6.7",
         "prettier": "3.0.0",
         "semantic-release-plugin-update-version-in-files": "^1.0.0",
         "ts-jest": "^29.0.0",
         "typescript": "^5.0.0"
       },
       "engines": {
-        "node": ">= 14"
+        "node": ">= 18"
       }
     },
     "node_modules/@ampproject/remapping": {
@@ -52,42 +50,42 @@
       }
     },
     "node_modules/@babel/code-frame": {
-      "version": "7.22.5",
-      "resolved": "https://registry.npmjs.org/@babel/code-frame/-/code-frame-7.22.5.tgz",
-      "integrity": "sha512-Xmwn266vad+6DAqEB2A6V/CcZVp62BbwVmcOJc2RPuwih1kw02TjQvWVWlcKGbBPd+8/0V5DEkOcizRGYsspYQ==",
+      "version": "7.21.4",
+      "resolved": "https://registry.npmjs.org/@babel/code-frame/-/code-frame-7.21.4.tgz",
+      "integrity": "sha512-LYvhNKfwWSPpocw8GI7gpK2nq3HSDuEPC/uSYaALSJu9xjsalaaYFOq0Pwt5KmVqwEbZlDu81aLXwBOmD/Fv9g==",
       "dev": true,
       "dependencies": {
-        "@babel/highlight": "^7.22.5"
+        "@babel/highlight": "^7.18.6"
       },
       "engines": {
         "node": ">=6.9.0"
       }
     },
     "node_modules/@babel/compat-data": {
-      "version": "7.22.5",
-      "resolved": "https://registry.npmjs.org/@babel/compat-data/-/compat-data-7.22.5.tgz",
-      "integrity": "sha512-4Jc/YuIaYqKnDDz892kPIledykKg12Aw1PYX5i/TY28anJtacvM1Rrr8wbieB9GfEJwlzqT0hUEao0CxEebiDA==",
+      "version": "7.21.7",
+      "resolved": "https://registry.npmjs.org/@babel/compat-data/-/compat-data-7.21.7.tgz",
+      "integrity": "sha512-KYMqFYTaenzMK4yUtf4EW9wc4N9ef80FsbMtkwool5zpwl4YrT1SdWYSTRcT94KO4hannogdS+LxY7L+arP3gA==",
       "dev": true,
       "engines": {
         "node": ">=6.9.0"
       }
     },
     "node_modules/@babel/core": {
-      "version": "7.22.5",
-      "resolved": "https://registry.npmjs.org/@babel/core/-/core-7.22.5.tgz",
-      "integrity": "sha512-SBuTAjg91A3eKOvD+bPEz3LlhHZRNu1nFOVts9lzDJTXshHTjII0BAtDS3Y2DAkdZdDKWVZGVwkDfc4Clxn1dg==",
+      "version": "7.21.8",
+      "resolved": "https://registry.npmjs.org/@babel/core/-/core-7.21.8.tgz",
+      "integrity": "sha512-YeM22Sondbo523Sz0+CirSPnbj9bG3P0CdHcBZdqUuaeOaYEFbOLoGU7lebvGP6P5J/WE9wOn7u7C4J9HvS1xQ==",
       "dev": true,
       "dependencies": {
         "@ampproject/remapping": "^2.2.0",
-        "@babel/code-frame": "^7.22.5",
-        "@babel/generator": "^7.22.5",
-        "@babel/helper-compilation-targets": "^7.22.5",
-        "@babel/helper-module-transforms": "^7.22.5",
-        "@babel/helpers": "^7.22.5",
-        "@babel/parser": "^7.22.5",
-        "@babel/template": "^7.22.5",
-        "@babel/traverse": "^7.22.5",
-        "@babel/types": "^7.22.5",
+        "@babel/code-frame": "^7.21.4",
+        "@babel/generator": "^7.21.5",
+        "@babel/helper-compilation-targets": "^7.21.5",
+        "@babel/helper-module-transforms": "^7.21.5",
+        "@babel/helpers": "^7.21.5",
+        "@babel/parser": "^7.21.8",
+        "@babel/template": "^7.20.7",
+        "@babel/traverse": "^7.21.5",
+        "@babel/types": "^7.21.5",
         "convert-source-map": "^1.7.0",
         "debug": "^4.1.0",
         "gensync": "^1.0.0-beta.2",
@@ -126,12 +124,12 @@
       "dev": true
     },
     "node_modules/@babel/generator": {
-      "version": "7.22.5",
-      "resolved": "https://registry.npmjs.org/@babel/generator/-/generator-7.22.5.tgz",
-      "integrity": "sha512-+lcUbnTRhd0jOewtFSedLyiPsD5tswKkbgcezOqqWFUVNEwoUTlpPOBmvhG7OXWLR4jMdv0czPGH5XbflnD1EA==",
+      "version": "7.21.5",
+      "resolved": "https://registry.npmjs.org/@babel/generator/-/generator-7.21.5.tgz",
+      "integrity": "sha512-SrKK/sRv8GesIW1bDagf9cCG38IOMYZusoe1dfg0D8aiUe3Amvoj1QtjTPAWcfrZFvIwlleLb0gxzQidL9w14w==",
       "dev": true,
       "dependencies": {
-        "@babel/types": "^7.22.5",
+        "@babel/types": "^7.21.5",
         "@jridgewell/gen-mapping": "^0.3.2",
         "@jridgewell/trace-mapping": "^0.3.17",
         "jsesc": "^2.5.1"
@@ -141,13 +139,13 @@
       }
     },
     "node_modules/@babel/helper-compilation-targets": {
-      "version": "7.22.5",
-      "resolved": "https://registry.npmjs.org/@babel/helper-compilation-targets/-/helper-compilation-targets-7.22.5.tgz",
-      "integrity": "sha512-Ji+ywpHeuqxB8WDxraCiqR0xfhYjiDE/e6k7FuIaANnoOFxAHskHChz4vA1mJC9Lbm01s1PVAGhQY4FUKSkGZw==",
+      "version": "7.21.5",
+      "resolved": "https://registry.npmjs.org/@babel/helper-compilation-targets/-/helper-compilation-targets-7.21.5.tgz",
+      "integrity": "sha512-1RkbFGUKex4lvsB9yhIfWltJM5cZKUftB2eNajaDv3dCMEp49iBG0K14uH8NnX9IPux2+mK7JGEOB0jn48/J6w==",
       "dev": true,
       "dependencies": {
-        "@babel/compat-data": "^7.22.5",
-        "@babel/helper-validator-option": "^7.22.5",
+        "@babel/compat-data": "^7.21.5",
+        "@babel/helper-validator-option": "^7.21.0",
         "browserslist": "^4.21.3",
         "lru-cache": "^5.1.1",
         "semver": "^6.3.0"
@@ -169,151 +167,151 @@
       }
     },
     "node_modules/@babel/helper-environment-visitor": {
-      "version": "7.22.5",
-      "resolved": "https://registry.npmjs.org/@babel/helper-environment-visitor/-/helper-environment-visitor-7.22.5.tgz",
-      "integrity": "sha512-XGmhECfVA/5sAt+H+xpSg0mfrHq6FzNr9Oxh7PSEBBRUb/mL7Kz3NICXb194rCqAEdxkhPT1a88teizAFyvk8Q==",
+      "version": "7.21.5",
+      "resolved": "https://registry.npmjs.org/@babel/helper-environment-visitor/-/helper-environment-visitor-7.21.5.tgz",
+      "integrity": "sha512-IYl4gZ3ETsWocUWgsFZLM5i1BYx9SoemminVEXadgLBa9TdeorzgLKm8wWLA6J1N/kT3Kch8XIk1laNzYoHKvQ==",
       "dev": true,
       "engines": {
         "node": ">=6.9.0"
       }
     },
     "node_modules/@babel/helper-function-name": {
-      "version": "7.22.5",
-      "resolved": "https://registry.npmjs.org/@babel/helper-function-name/-/helper-function-name-7.22.5.tgz",
-      "integrity": "sha512-wtHSq6jMRE3uF2otvfuD3DIvVhOsSNshQl0Qrd7qC9oQJzHvOL4qQXlQn2916+CXGywIjpGuIkoyZRRxHPiNQQ==",
+      "version": "7.21.0",
+      "resolved": "https://registry.npmjs.org/@babel/helper-function-name/-/helper-function-name-7.21.0.tgz",
+      "integrity": "sha512-HfK1aMRanKHpxemaY2gqBmL04iAPOPRj7DxtNbiDOrJK+gdwkiNRVpCpUJYbUT+aZyemKN8brqTOxzCaG6ExRg==",
       "dev": true,
       "dependencies": {
-        "@babel/template": "^7.22.5",
-        "@babel/types": "^7.22.5"
+        "@babel/template": "^7.20.7",
+        "@babel/types": "^7.21.0"
       },
       "engines": {
         "node": ">=6.9.0"
       }
     },
     "node_modules/@babel/helper-hoist-variables": {
-      "version": "7.22.5",
-      "resolved": "https://registry.npmjs.org/@babel/helper-hoist-variables/-/helper-hoist-variables-7.22.5.tgz",
-      "integrity": "sha512-wGjk9QZVzvknA6yKIUURb8zY3grXCcOZt+/7Wcy8O2uctxhplmUPkOdlgoNhmdVee2c92JXbf1xpMtVNbfoxRw==",
+      "version": "7.18.6",
+      "resolved": "https://registry.npmjs.org/@babel/helper-hoist-variables/-/helper-hoist-variables-7.18.6.tgz",
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-        "@babel/types": "^7.22.5"
+        "@babel/types": "^7.18.6"
       },
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         "node": ">=6.9.0"
       }
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+      "resolved": "https://registry.npmjs.org/@babel/helper-module-imports/-/helper-module-imports-7.21.4.tgz",
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+        "@babel/types": "^7.21.4"
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       }
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-      "version": "3.0.1",
-      "resolved": "https://registry.npmjs.org/form-data/-/form-data-3.0.1.tgz",
-      "integrity": "sha512-RHkBKtLWUVwd7SqRIvCZMEvAMoGUp0XU+seQiZejj0COz3RI3hWP4sCv3gZWWLjJTd7rGwcsF5eKZGii0r/hbg==",
-      "dev": true,
-      "dependencies": {
-        "asynckit": "^0.4.0",
-        "combined-stream": "^1.0.8",
-        "mime-types": "^2.1.12"
-      },
-      "engines": {
-        "node": ">= 6"
-      }
-    },
     "node_modules/forwarded": {
       "version": "0.2.0",
       "resolved": "https://registry.npmjs.org/forwarded/-/forwarded-0.2.0.tgz",
@@ -3411,25 +3375,6 @@
         "node": ">= 0.6"
       }
     },
-    "node_modules/fromentries": {
-      "version": "1.3.2",
-      "resolved": "https://registry.npmjs.org/fromentries/-/fromentries-1.3.2.tgz",
-      "integrity": "sha512-cHEpEQHUg0f8XdtZCc2ZAhrHzKzT0MrFUTcvx+hfxYu7rGMDc5SKoXFh+n4YigxsHXRzc6OrCshdR1bWH6HHyg==",
-      "funding": [
-        {
-          "type": "github",
-          "url": "https://github.com/sponsors/feross"
-        },
-        {
-          "type": "patreon",
-          "url": "https://www.patreon.com/feross"
-        },
-        {
-          "type": "consulting",
-          "url": "https://feross.org/support"
-        }
-      ]
-    },
     "node_modules/fs.realpath": {
       "version": "1.0.0",
       "resolved": "https://registry.npmjs.org/fs.realpath/-/fs.realpath-1.0.0.tgz",
@@ -5749,9 +5694,10 @@
       "dev": true
     },
     "node_modules/lru-cache": {
-      "version": "9.1.2",
-      "resolved": "https://registry.npmjs.org/lru-cache/-/lru-cache-9.1.2.tgz",
-      "integrity": "sha512-ERJq3FOzJTxBbFjZ7iDs+NiK4VI9Wz+RdrrAB8dio1oV+YvdPzUEE4QNiT2VD51DkIbCYRUUzCRkssXCHqSnKQ==",
+      "version": "9.1.1",
+      "resolved": "https://registry.npmjs.org/lru-cache/-/lru-cache-9.1.1.tgz",
+      "integrity": "sha512-65/Jky17UwSb0BuB9V+MyDpsOtXKmYwzhyl+cOa9XUiI4uV2Ouy/2voFP3+al0BjZbJgMBD8FojMpAf+Z+qn4A==",
+      "dev": true,
       "engines": {
         "node": "14 || >=16.14"
       }
@@ -5926,6 +5872,9 @@
       "version": "2.6.11",
       "resolved": "https://registry.npmjs.org/node-fetch/-/node-fetch-2.6.11.tgz",
       "integrity": "sha512-4I6pdBY1EthSqDmJkiNk3JIT8cswwR9nfeW/cPdUagJYEQG7R95WRH74wpz7ma8Gh/9dI9FP+OU+0E4FvtA55w==",
+      "dev": true,
+      "optional": true,
+      "peer": true,
       "dependencies": {
         "whatwg-url": "^5.0.0"
       },
@@ -5944,17 +5893,26 @@
     "node_modules/node-fetch/node_modules/tr46": {
       "version": "0.0.3",
       "resolved": "https://registry.npmjs.org/tr46/-/tr46-0.0.3.tgz",
-      "integrity": "sha512-N3WMsuqV66lT30CrXNbEjx4GEwlow3v6rr4mCcv6prnfwhS01rkgyFdjPNBYd9br7LpXV1+Emh01fHnq2Gdgrw=="
+      "integrity": "sha512-N3WMsuqV66lT30CrXNbEjx4GEwlow3v6rr4mCcv6prnfwhS01rkgyFdjPNBYd9br7LpXV1+Emh01fHnq2Gdgrw==",
+      "dev": true,
+      "optional": true,
+      "peer": true
     },
     "node_modules/node-fetch/node_modules/webidl-conversions": {
       "version": "3.0.1",
       "resolved": "https://registry.npmjs.org/webidl-conversions/-/webidl-conversions-3.0.1.tgz",
-      "integrity": "sha512-2JAn3z8AR6rjK8Sm8orRC0h/bcl/DqL7tRPdGZ4I1CjdF+EaMLmYxBHyXuKL849eucPFhvBoxMsflfOb8kxaeQ=="
+      "integrity": "sha512-2JAn3z8AR6rjK8Sm8orRC0h/bcl/DqL7tRPdGZ4I1CjdF+EaMLmYxBHyXuKL849eucPFhvBoxMsflfOb8kxaeQ==",
+      "dev": true,
+      "optional": true,
+      "peer": true
     },
     "node_modules/node-fetch/node_modules/whatwg-url": {
       "version": "5.0.0",
       "resolved": "https://registry.npmjs.org/whatwg-url/-/whatwg-url-5.0.0.tgz",
       "integrity": "sha512-saE57nupxk6v3HY35+jzBwYa0rKSy0XR8JSxZPwgLr7ys0IBzhGviA1/TUGJLmSVqs8pb9AnvICXEuOHLprYTw==",
+      "dev": true,
+      "optional": true,
+      "peer": true,
       "dependencies": {
         "tr46": "~0.0.3",
         "webidl-conversions": "^3.0.0"
@@ -5967,9 +5925,9 @@
       "dev": true
     },
     "node_modules/node-releases": {
-      "version": "2.0.12",
-      "resolved": "https://registry.npmjs.org/node-releases/-/node-releases-2.0.12.tgz",
-      "integrity": "sha512-QzsYKWhXTWx8h1kIvqfnC++o0pEmpRQA/aenALsL2F4pqNVr7YzcdMlDij5WBnwftRbJCNJL/O7zdKaxKPHqgQ==",
+      "version": "2.0.13",
+      "resolved": "https://registry.npmjs.org/node-releases/-/node-releases-2.0.13.tgz",
+      "integrity": "sha512-uYr7J37ae/ORWdZeQ1xxMJe3NtdmqMC/JZK+geofDrkLUApKRHPd18/TxtBOJ4A0/+uUIliorNrfYV6s1b02eQ==",
       "dev": true
     },
     "node_modules/normalize-path": {
@@ -6789,9 +6747,9 @@
       }
     },
     "node_modules/strip-ansi": {
-      "version": "7.1.0",
-      "resolved": "https://registry.npmjs.org/strip-ansi/-/strip-ansi-7.1.0.tgz",
-      "integrity": "sha512-iq6eVVI64nQQTRYq2KtEg2d2uU7LElhTJwsH4YzIHZshxlgZms/wIc4VoDQTlG/IvVIrBKG06CrZnp0qv7hkcQ==",
+      "version": "7.0.1",
+      "resolved": "https://registry.npmjs.org/strip-ansi/-/strip-ansi-7.0.1.tgz",
+      "integrity": "sha512-cXNxvT8dFNRVfhVME3JAe98mkXDYN2O1l7jmcwMnOslDeESg1rF/OZMtK0nRAhiari1unG5cD4jG3rapUAkLbw==",
       "dev": true,
       "dependencies": {
         "ansi-regex": "^6.0.1"
@@ -7091,16 +7049,16 @@
       }
     },
     "node_modules/typescript": {
-      "version": "5.1.3",
-      "resolved": "https://registry.npmjs.org/typescript/-/typescript-5.1.3.tgz",
-      "integrity": "sha512-XH627E9vkeqhlZFQuL+UsyAXEnibT0kWR2FWONlr4sTjvxyJYnyefgrkyECLzM5NenmKzRAy2rR/OlYLA1HkZw==",
+      "version": "5.0.4",
+      "resolved": "https://registry.npmjs.org/typescript/-/typescript-5.0.4.tgz",
+      "integrity": "sha512-cW9T5W9xY37cc+jfEnaUvX91foxtHkza3Nw3wkoF4sSlKn0MONdkdEndig/qPBWXNkmplh3NzayQzCiHM4/hqw==",
       "dev": true,
       "bin": {
         "tsc": "bin/tsc",
         "tsserver": "bin/tsserver"
       },
       "engines": {
-        "node": ">=14.17"
+        "node": ">=12.20"
       }
     },
     "node_modules/universal-github-app-jwt": {
diff --git a/package.json b/package.json
index db110c1ac..39b0f90d7 100644
--- a/package.json
+++ b/package.json
@@ -18,33 +18,31 @@
   "author": "Gregor Martynus (https://github.com/gr2m)",
   "license": "MIT",
   "dependencies": {
-    "@octokit/auth-app": "^4.0.13",
-    "@octokit/auth-unauthenticated": "^3.0.0",
-    "@octokit/core": "^4.0.0",
-    "@octokit/oauth-app": "^4.0.7",
-    "@octokit/plugin-paginate-rest": "^6.0.0",
-    "@octokit/types": "^9.0.0",
-    "@octokit/webhooks": "^10.0.0"
+    "@octokit/auth-app": "^6.0.0",
+    "@octokit/auth-unauthenticated": "^5.0.0",
+    "@octokit/core": "^5.0.0",
+    "@octokit/oauth-app": "^6.0.0",
+    "@octokit/plugin-paginate-rest": "^8.0.0",
+    "@octokit/types": "^11.1.0",
+    "@octokit/webhooks": "^12.0.1"
   },
   "devDependencies": {
     "@octokit/tsconfig": "^2.0.0",
     "@types/jest": "^29.0.0",
     "@types/node": "^18.0.0",
-    "@types/node-fetch": "^2.5.8",
     "esbuild": "^0.18.0",
     "express": "^4.17.1",
     "fetch-mock": "^9.10.7",
     "glob": "^10.0.0",
     "jest": "^29.0.0",
     "mockdate": "^3.0.2",
-    "node-fetch": "^2.6.7",
     "prettier": "3.0.0",
     "semantic-release-plugin-update-version-in-files": "^1.0.0",
     "ts-jest": "^29.0.0",
     "typescript": "^5.0.0"
   },
   "engines": {
-    "node": ">= 14"
+    "node": ">= 18"
   },
   "jest": {
     "transform": {
diff --git a/src/middleware/node/index.ts b/src/middleware/node/index.ts
index cbe6054e1..f48e50763 100644
--- a/src/middleware/node/index.ts
+++ b/src/middleware/node/index.ts
@@ -4,20 +4,19 @@
 type IncomingMessage = any;
 type ServerResponse = any;
 
-import { createNodeMiddleware as oauthNodeMiddleware } from "@octokit/oauth-app";
+import {
+  createNodeMiddleware as oauthNodeMiddleware,
+  sendNodeResponse,
+  unknownRouteResponse,
+} from "@octokit/oauth-app";
 import { createNodeMiddleware as webhooksNodeMiddleware } from "@octokit/webhooks";
 
 import { App } from "../../index";
-import { onUnhandledRequestDefault } from "./on-unhandled-request-default";
 import type { Options } from "../../types";
 
 export type MiddlewareOptions = {
   pathPrefix?: string;
   log?: Options["log"];
-  onUnhandledRequest?: (
-    request: IncomingMessage,
-    response: ServerResponse
-  ) => void;
 };
 
 function noop() {}
@@ -37,7 +36,6 @@ export function createNodeMiddleware(
   );
 
   const optionsWithDefaults = {
-    onUnhandledRequest: onUnhandledRequestDefault,
     pathPrefix: "/api/github",
     ...options,
     log,
@@ -46,41 +44,41 @@ export function createNodeMiddleware(
   const webhooksMiddleware = webhooksNodeMiddleware(app.webhooks, {
     path: optionsWithDefaults.pathPrefix + "/webhooks",
     log,
-    onUnhandledRequest: optionsWithDefaults.onUnhandledRequest,
   });
 
   const oauthMiddleware = oauthNodeMiddleware(app.oauth, {
     pathPrefix: optionsWithDefaults.pathPrefix + "/oauth",
-    onUnhandledRequest: optionsWithDefaults.onUnhandledRequest,
   });
 
-  return middleware.bind(null, optionsWithDefaults, {
+  return middleware.bind(
+    null,
+    optionsWithDefaults.pathPrefix,
     webhooksMiddleware,
-    oauthMiddleware,
-  });
+    oauthMiddleware
+  );
 }
 
 export async function middleware(
-  options: Required<MiddlewareOptions>,
-  { webhooksMiddleware, oauthMiddleware }: any,
+  pathPrefix: string,
+  webhooksMiddleware: any,
+  oauthMiddleware: any,
   request: IncomingMessage,
   response: ServerResponse,
   next?: Function
-) {
+): Promise<boolean> {
   const { pathname } = new URL(request.url as string, "http://localhost");
 
-  if (pathname === `${options.pathPrefix}/webhooks`) {
-    return webhooksMiddleware(request, response, next);
-  }
-  if (pathname.startsWith(`${options.pathPrefix}/oauth/`)) {
-    return oauthMiddleware(request, response, next);
+  if (pathname.startsWith(`${pathPrefix}/`)) {
+    if (pathname === `${pathPrefix}/webhooks`) {
+      webhooksMiddleware(request, response);
+    } else if (pathname.startsWith(`${pathPrefix}/oauth/`)) {
+      oauthMiddleware(request, response);
+    } else {
+      sendNodeResponse(unknownRouteResponse(request), response);
+    }
+    return true;
+  } else {
+    next?.();
+    return false;
   }
-
-  const isExpressMiddleware = typeof next === "function";
-  if (isExpressMiddleware) {
-    // @ts-ignore `next` must be a function as we check two lines above
-    return next();
-  }
-
-  return options.onUnhandledRequest(request, response);
 }
diff --git a/src/middleware/node/on-unhandled-request-default.ts b/src/middleware/node/on-unhandled-request-default.ts
deleted file mode 100644
index 3664190fc..000000000
--- a/src/middleware/node/on-unhandled-request-default.ts
+++ /dev/null
@@ -1,19 +0,0 @@
-// remove type imports from http for Deno compatibility
-// see https://github.com/octokit/octokit.js/issues/24#issuecomment-817361886
-// import { IncomingMessage, ServerResponse } from "http";
-type IncomingMessage = any;
-type ServerResponse = any;
-
-export function onUnhandledRequestDefault(
-  request: IncomingMessage,
-  response: ServerResponse
-) {
-  response.writeHead(404, {
-    "content-type": "application/json",
-  });
-  response.end(
-    JSON.stringify({
-      error: `Unknown route: ${request.method} ${request.url}`,
-    })
-  );
-}
diff --git a/test/node-middleware.test.ts b/test/node-middleware.test.ts
index f3378cc8b..ef21abd73 100644
--- a/test/node-middleware.test.ts
+++ b/test/node-middleware.test.ts
@@ -1,5 +1,4 @@
 import { createServer } from "http";
-import fetch from "node-fetch";
 
 // import without types
 const express = require("express");
@@ -36,30 +35,99 @@ x//0u+zd/R/QRUzLOw4N72/Hu+UG6MNt5iDZFCtapRaKt6OvSBwy8w==
 -----END RSA PRIVATE KEY-----`;
 
 describe("createNodeMiddleware()", () => {
-  test("unknown route", async () => {
-    const app = new App({
-      appId: APP_ID,
-      privateKey: PRIVATE_KEY,
-      webhooks: {
-        secret: "mysecret",
-      },
-      oauth: {
-        clientId: "",
-        clientSecret: "",
-      },
+  test("unknown route (outside pathPrefix)", async () => {
+    const middleware = createNodeMiddleware({} as unknown as App);
+    const server = createServer(async (req, res) => {
+      if (!(await middleware(req, res))) {
+        res.writeHead(404, { "Content-Type": "text/plain" });
+        res.write("Not found.");
+        res.end();
+      }
+    }).listen();
+    // @ts-expect-error complains about { port } although it's included in returned AddressInfo interface
+    const { port } = server.address();
+
+    const response = await fetch(`http://localhost:${port}/unknown-route`);
+
+    expect(response.status).toEqual(404);
+    expect(response.headers.get("Content-Type")).toBe("text/plain");
+    await expect(response.text()).resolves.toBe("Not found.");
+
+    server.close();
+  });
+
+  test("unknown route (within pathPrefix)", async () => {
+    const middleware = createNodeMiddleware({} as unknown as App);
+    const server = createServer(async (req, res) => {
+      if (!(await middleware(req, res))) {
+        res.writeHead(404, { "Content-Type": "text/plain" });
+        res.write("Not found.");
+        res.end();
+      }
+    }).listen();
+    // @ts-expect-error complains about { port } although it's included in returned AddressInfo interface
+    const { port } = server.address();
+
+    const response = await fetch(
+      `http://localhost:${port}/api/github/unknown-route`,
+    );
+
+    expect(response.status).toEqual(404);
+    expect(response.headers.get("Content-Type")).toBe("application/json");
+    await expect(response.json()).resolves.toEqual({
+      error: "Unknown route: GET /api/github/unknown-route",
     });
 
-    const server = createServer(createNodeMiddleware(app)).listen();
+    server.close();
+  });
+
+  test("unknown route (within webhooks path)", async () => {
+    const middleware = createNodeMiddleware({} as unknown as App);
+    const server = createServer(async (req, res) => {
+      if (!(await middleware(req, res))) {
+        res.writeHead(404, { "Content-Type": "text/plain" });
+        res.write("Not found.");
+        res.end();
+      }
+    }).listen();
     // @ts-expect-error complains about { port } although it's included in returned AddressInfo interface
     const { port } = server.address();
 
-    const response = await fetch(`http://localhost:${port}/unknown-route`);
+    const response = await fetch(
+      `http://localhost:${port}/api/github/webhooks`,
+    );
 
     expect(response.status).toEqual(404);
-    await expect(response.text()).resolves.toMatch(
-      /Unknown route: GET \/unknown-route/,
+    expect(response.headers.get("Content-Type")).toBe("application/json");
+    await expect(response.json()).resolves.toEqual({
+      error: "Unknown route: GET /api/github/webhooks",
+    });
+
+    server.close();
+  });
+
+  test("unknown route (within oauth pathPrefix)", async () => {
+    const middleware = createNodeMiddleware({} as unknown as App);
+    const server = createServer(async (req, res) => {
+      if (!(await middleware(req, res))) {
+        res.writeHead(404, { "Content-Type": "text/plain" });
+        res.write("Not found.");
+        res.end();
+      }
+    }).listen();
+    // @ts-expect-error complains about { port } although it's included in returned AddressInfo interface
+    const { port } = server.address();
+
+    const response = await fetch(
+      `http://localhost:${port}/api/github/oauth/unknown-route`,
     );
 
+    expect(response.status).toEqual(404);
+    expect(response.headers.get("Content-Type")).toBe("application/json");
+    await expect(response.json()).resolves.toEqual({
+      error: "Unknown route: GET /api/github/oauth/unknown-route",
+    });
+
     server.close();
   });
 
diff --git a/test/readme-examples.test.ts b/test/readme-examples.test.ts
index 757e9ec83..7ad3f9f40 100644
--- a/test/readme-examples.test.ts
+++ b/test/readme-examples.test.ts
@@ -219,7 +219,7 @@ describe("README examples", () => {
     // @ts-ignore
     const { port } = server.address();
     const url = `http://localhost:${port}/api/github/webhooks`;
-    const data = {
+    const data = JSON.stringify({
       action: "opened",
       installation: {
         id: 123,
@@ -233,7 +233,7 @@ describe("README examples", () => {
       issue: {
         number: 456,
       },
-    };
+    });
 
     await request({
       method: "POST",